DINOSAURICON MN


M

Afbeeldingen van Machairasaurus  <–

 

Malawisaurus (Greek for “Malawi lizard”);
Habitat:Woodlands of Africa /Early Cretaceous (125-115 million years ago)/Large size; armor plating on back

(4.3 meters) (9.1 meters) (10,900 kg)

Malawi, Africa. Discovered by Dr. Louis Jacobs, Malawisaurus closely resembles Andesaurus from South America. Its discovery provided a missing link in the evolutionary story of titanosaurids. It is the only Titanosaurid for which skull material (but not a whole skull) is known.

Moreso than the still-mysterious Titanosaurus, Malawisaurus can arguably be considered the “type specimen” for titanosaurs, the lightly armored descendants of the giant sauropods of the Jurassic period. Malawisaurus is one of the few titanosaurs for which we have direct evidence of a skull (albeit only a partial one that includes most of the upper and lower jaw), and fossilized scutes have been found in the vicinity of its remains, evidence of the armor plating that once lined this herbivore’s neck and back.

( Incidentally, Malawisaurus was once considered a species of the now-invalid genus Gigantosaurus–not to be confused with Giganotosaurus (note that extra “o”), which wasn’t a titanosaur at all but a large theropod.

Malawisaurus dixeyi is one of the most complete titanosaur sauropods described in the scientific literature to date.
It is known from skull material (which is a rare case for sauropods in general, not to mention titanosaurs), most of the cervical (neck) vertebrae, all the dorsal (back) vertebrae, all the sacral vertebrae and a large portion of the caudal (tail)
vertebrae as well as most of the appendicular elements excluding the ilia, pubis and scapula
Malawisaurus was rather small (for a sauropod). The projection of its length was about 11.05 meters (that is what the apparent length is from snout to the tip of the tail in the top view of the skeletal).
However, along the vertebral series it was about 11.25 meters long. This is in comparison to some titanosaurs which reached over 35 meters in length, and at least one poorly known diplodocid sauropod reached a length of 60-70 meters long. It massed around 2.52-2.59 tonnes, compared to the largest titanosaurs which may have massed up to 100 tonnes and the largest sauropod ever, Amphicoelias fragillimus, probably massed close to 200 tonnes.
Malawisaurus is even exceeded in mass by a number of living terrestrial mammal species, as Loxodonta africana (African Bush Elephant), L. cyclotis (African Forest Elephant), Elephas maximus (Asian Elephant), Hippopotamus amphibius (Hippo), Ceratotherium simum (White Rhino), Rhinoceros unicornis (Indian Rhino) all of which have members that well exceed 2,500 kg in mass.
The largely aquatic (although they breed, fight and rest on land) species in the genus Mirounga (the elephant seals) also can mass well over 2,500 kg.
Notes on the reconstruction by
The gray bones indicate bones and parts of bones that aren’t preserved and so are based off of related species.
The white bones indicate preserved bones and the pale yellowish colored bones are bones that are preserved but not figured so their shape is conjectural.
Finally, the light blue bones are bones that are from another specimen to help fill in some gaps of knowledge in the caudal series.
In comparison to earlier reconstructions some updatring changes were made :
First, the metacarpal configuration of the forelimb was changed;
second, the black outline around the forelimb was changed to reflect a more robust musculature;
similarly, the black outline around the lower part of the hindlimb was changed to reflect a more robust musculature; also, the size of the ischium is now larger because it was incorrectly scaled in the earlier version;
Palaeozoologist also  changed the length of the pubis to be more similar proportionally to that of Futalognkosaurus with regards the change of the size of the ischium;

finally, the black outline of the belly has been extended and the outline around the anterior to mid-caudals has been extended laterally to reflect new thoughts about the tail musculature in dinosaurs.

Dentaries and teeth demonstrate the presence of at least two sauropods in Malawi,Malawisaurus and Karongasaurus.

Based on differences in the morphology of the caudal vertebrae that have been generally recognized as significant, titanosaurian caudal vertebrae from Malawi also represent at least two taxa including Malawisaurus and Titanosauridae

indet. Malawisaurus has platycoelous medial caudal vertebrae, whereas a procoelous middle or posterior caudal vertebra is distinct from Malawisaurus and referred to Titanosauridae indet. A third taxon may be represented by vertebra referred to Titanosauria indet.

Phylogenetic analyses that include Aeolosaurus, Alamosaurus, Andesaurus, Antarctosaurus, Argentinosaurus, Epachthosaurus, Malawisaurus, Nemegtosaurus, Neuquensaurus,Opisthocoelicaudia, Quaesitosaurus, Rapetosaurus, Saltasaurus, and Titanosaurus (Upchurch 1995,1998,1999;Salgado et al. 1997;Curry Rogers and Forster 2001; see also Wilson 2002) indicate that Andesaurus and Malawisaurus are basal titanosaurians.

In fact, Malawisaurus is the most complete Early Cretaceous titanosaurian known.

It is represented by cranial elements, 18 cervical, 10 dorsal, six sacral, and 51 caudal vertebrae, 24 chevrons, pectoral elements, pelvic elements, and dermal armor. Phylogenetic analyses also indicate that taxa with cylindrical teeth and strongly procoelous posterior caudal vertebrae (or opisthocoelous in the case of Opisthocoelicaudia) are more derived than those with broad teeth and platycoelous middle and distal caudal vertebrae.

Skull Shape and Morphological Diversity in Malawi Sauropods. Cranial material attributed to titanosaurians includes one or two partial braincases of Titanosaurus indicus from the Late Cretaceous of India (Berman and Jain 1982;Chatterjee and Rudra 1996), a maxilla from India (von Huene and Matley 1933), a partial braincase and partial skull roof of Saltasaurus (PVL 4017-161) from the Late Cretaceous of Patagonia (Powell 1986,2003), a premaxilla of Titanosauridae indet. from the Late Cretaceous of Patagonia (Scuitto and Martinez 1994), a premaxilla (PVL 3670-12) that Powell (1979) identified as Laplatasaurus but later referred to as Titanosauridae indet. (Powell 1986,2003), a fragmentary braincase from the Late Cretaceous of France (Le Loeuff et al. 1989), the braincase, quadrate, quadratojugal, squamosal, and the lower jaws of Antarctosaurus wichmannianus (MACN 6904) from the Late Cretaceous of Patagonia (von Huene 1929;Powell 1986,2003), two partial braincases of Antarctosaurus septentrionalis from the Late Cretaceous of India (von Huene and Matley 1933;Chatterjee and Rudra 1996), a nearly complete disarticulated cranium of Rapetosaurus from the Late Cretaceous of Madagascar (Curry Rogers and Forster 2001,2004), and the specimens of Malawisaurus from the Early Cretaceous of Malawi.

There are two basic morphs of sauropod skulls: one high and short, the other low and elongate (Wilson 2002). The high, short morph is referred to as a macronarian skull, whereas the low, elongate morph is generally referred to as a diplodocoid skull (Coombs 1975;McIntosh and Berman 1975;Berman and McIntosh 1978;Salgado and Calvo 1997;Wilson 2002). The macronarian skull is present in Brachiosaurus, Camarasaurus, Datousaurus, Euhelopus, Mamenchisaurus, Omeisaurus, Shunosaurus, and most prosauropods, whereas the diplodocoid skull is present in Apatosaurus, Diplodocus, and dicraeosaurids (Salgado and Calvo 1997; see also Tidwell and Carpenter, 2003).

Comparison of cranial features of Malawisaurus with macronarian and diplodocoidskulls suggests that Malawisaurus had a high, short macronarian skull (Figure 31). The anterior section of the suture between premaxilla and maxilla appears to have been nearly vertical as suggested by the highly angled articular surface for the maxilla on the premaxilla. The high premaxilla also suggests that Malawisaurus had a high, short, and blunt snout, and separate nares positioned rostrally and facing laterally. The tooth row extends more than half the length of the dentary. The mandibular symphysis is oblique to the long axis of the mandible. The occipital condyle projected posteroventrally and the basipterygoid processes projected ventrally. The quadrate axis was nearly vertical, the pterygoid process of the quadrate was approximately perpendicular to the long axis of the pterygoid, the pterygoid process was directed anteriorly as in Brachiosaurus (Janensch 1935) and Camarasaurus (Osborn and Mook 1921), and the mandibular articulation was placed posteriorly beneath the level of the occipital condyle. Thus, this study demonstrates that at least some titanosaurians, including Malawisaurus, had high and short crania, as compared to others such as Rapetosaurus, which had low and elongate crania (Curry Rogers and Forster 2001,2004).

In addition, the titanosaurians Nemegtosaurus (Nowinski 1971), and Quaesitosaurus (Kurzanov and Bannikov 1983; Curry Rogers and Forster 2001) had slender teeth restricted to the anterior portion of the mandible, and also had low and elongate crania. If that association of characters is general within titanosaurians, by implication Karongasaurus would also have had a low and elongate cranium. Further, phylogenetic analyses indicate that titanosaurians with slender teeth and which have strongly procoelous middle and posterior caudal vertebrae are derived relative to basal titanosaurians (Upchurch 1995;Salgado et al. 1997;Curry Rogers and Forster 2001). Thus, both Karongasaurus and Titanosauridae indet. from the Dinosaur Beds are derived relative toMalawisaurus. If those characters are linked, the vertebra assigned to Titanosauridae indet. may belong to Karongasaurus.

In any case, Malawisaurus and Karongasaurus are two distinct titanosaurian taxa that coexisted in the Early Cretaceous of Malawi and exhibited quite different morphological features, certainly in their lower jaws and teeth, and probably also in their skull shapes. Cylindrical teeth, an anteriorly restricted tooth row, and a long, low skull shape evolved as a complex at least twice, once within diplodocoids and once within titanosaurians. This character complex is functionally and adaptively important for feeding. Its multiple origins, and its variance from the macronarian skull pattern, implies that Malawisaurus and Karongasaurus were ecologically distinct. If so, differences seen in the lower jaw, teeth, and probably the skull of these herbivores were significant in the ecological partitioning of their Early Cretaceous environment. Although the macronarian and diplodocoid skull morphs are well known to occur together, for example in the Jurassic of Africa, in the Early Cretaceous of Malawi approximately equally divergent skull morphs are exhibited at a lower systematic level among titanosaurians alone

 

http://www.paleofile.com/Dinosaurs/Armor/Maleevus.asp

 

maleevus

 

Sketch of the sauropod Mamenchisaurus.

“Grootste dinoskelet” tentoongesteld in Japan

Het Gunma Museum of Natural History in Japan meent het grootste skelet van een dinosaurus in zijn collectie te hebben. Het gaat om een replica van een volledig skelet van een 35 meter lange Mamenchisaurus, een planteneter. Het skelet werd gevonden in de Gobiwoestijn in China in 2001. (ruwe beelden APTN – 13/07/09)

http://www.deredactie.be/cm/vrtnieuws/videozone/archief/nieuws/buitenland/1.562835

Click here to view larger image.

  Click to view animal family tree

Mamenchisaurus                                                                                                    //   145–160 million years ago, //Late Jurassic

compared to an African elephant and a woman

http://museumvictoria.com.au/melbournemuseum/discoverycentre/dinosaur-walk/meet-the-skeletons/mamenchisaurus/

 
Groep Maniraptorae

Bambiraptor
Bambiraptor. Photo © Rob Gay.

Bird’s closests relatives : Maniraptora ( “seizing hands ” )

We all know now that birds are dinosaurs, right? The clade Maniraptora (which is defined as containing all dinosaurs closer to birds than to ornithomimids ) is the group of theropod dinosaurs that many paleontologists believe birds were derived from some 150 or so million years ago, in the Jurassic period. Hence, according to phylogenetic taxonomy, birds are by definition maniraptorans, and the other maniraptorans are their closest relatives.

The exact membership of this clade is highly disputed; any coelurosaur could be quite convincingly argued to be a member of Maniraptora. There is much convergent evolution apparent in

Velociraptor skull
Velociraptor skull. Photo © Rob Gay.
We’ll present some of the most accepted members of the clade Maniraptora; their position may very well change with future studies.Maniraptora, which makes the resolution of their phylogeny difficult; a problem compounded by their poor fossil record. Even without considering birds, Maniraptora is a diverse and interesting group, with many specimens that outwardly look dissimilar, but have enough structures in common (synapomorphies) to unite them as a group. Maniraptorans are united by the possession of modified elements in the wrist; the semilunate carpal is a bone unique to this group — along with other modifications of the forelimb, it makes the flight stroke in birds possible, and was probably co-opted by birds for flight from a grasping function. Other characteristics present in typical maniraptorans include a fused clavicle (furcula, or “collar bone”) and sternum (“breast bone”), a pubis (part of the pelvis) that points downwards rather than forwards as in typical saurischians, a shortened and distally stiffened tail, long arms, and a manus (hand) which is larger than the pes (foot).

 

Major Maniraptoran groups:

Aves: The birds. Dromaeosaurs: The “raptor” dinosaurs. Troodontids: The smartest non-avian dinosaurs? Therizinosaurs: Bizarre plant-eating theropods? Oviraptors: Strange maniraptorans with evidence of devoted parental care
feather chart
This phylogenetic tree represents the evolutionary view of dinosaur-to-bird evolution. The green dot is the proposed branch point for theropods.
The yellow dot—placed after ornithomimids are said to have branched off—represents the beginning of maniraptorans.
Evolutionists generally believe that maniraptorans evolved into birds.
The current paleontological findings associated with each group—filaments, feathers, and wings—are indicated. Note that the ornithomimids are noted to have all three, even though the fossilized evidence only consists of some marks on bony surfaces without any actual feathers or wings being present.
Also note that the groupings include actual birds, some being extinct. 13 Image published in D. Zelenitsky et al.,10 through www.sciencemag.org.
.

GROEP : Marginocephalia *Marginocephalia

 

Afbeeldingen van Masiakasaurus knofleri  <—

http://nl.pinterest.com/tsjok/dinosauricon-a-abelisauridae/

Masiakasaurus knofleri - newly discovered dinosaur from Madagascar

http://www.dinosaur-world.com/weird_dinosaurs/masiakasaurus_knofleri.htm

 

Eerst kruipen, dan rennen

juli 2005

Zes prachtig geconserveerde dino-eieren werden in 1978 opgegraven in het Golden Gate Highlands National Park in Zuid Afrika.

De eieren waren zó fragiel, dat het 25 jaar zou duren voordat twee ervan werden opengemaakt. Onderzoekster Diana Scott is een jaar in de weer geweest met tandartsboortjes en andere piepkleine instrumenten om twee van de embryo’s in de zes centimeter grote eieren vrij te prepareren. Het zijn de oudste fossiele dino-embryo’s die ooit zijn gevonden. De eieren zijn afkomstig van Massospondylus carinatus, een vegetarische dino uit het late Trias en het vroege Jura, zo’n 220 tot 183 miljoen jaar geleden.

Een volwassen exemplaar mat vijf meter van staart tot kop, en liep rechtop op zijn achterpoten. De vorm van het hoofd, de nek en de bovenste ledematen van het embryo lijkt er echter op te wijzen dat de jonge exemplaren van de soort eerst een tijd op vier ledematen in het rond kropen, vooraleer ze op twee achterpoten gingen lopen. Vooral het grote hoofd was opvallend: dat moet veel te zwaar zijn geweest voor de lange nek. Ook opvallend was het feit dat de embryo’s geen tanden hebben. Dat betekent dat ze van hun ouders afhankelijk waren voor zorg en eten. Volgens Robert Reisz, een van de onderzoekers, is die bevinding – als ze juist is – de alleroudste aanwijzing van ouderlijke zorg

Cast of a Muttaburrasaurus skeleton.

montanoceratops-old-990x404Brown and Schlaikjer’s reconstruction of what we now call Montanoceratops, with the cheek bone confused for a nose horn. From Brown and Schlaikjer, 1942.
A restoration on Montanoceratops as we know the dinosaur now. Art by Nobu Tamura, image from Wikipedia.Chinnery, B., Weishamphel, D. 1998. Montanoceratops cerorhynchus (Dinosauria: Ceratopsia) and relationships among basal neoceratopsians. Journal of Vertebrate Paleontology. 18, 3: 569-585Makovicky, P. 2010. A redescription of the Montanoceratops cerorhynchus holotype with a review of referred material, pp. 68-82, in Ryan, M., Chinnery-Allgeier, B., Eberth, D., eds., New Perspectives on Horned Dinosaurs. Bloomington: Indiana University Press.

N

Nasuroceratops 

Een dinosaurus met een “grote neus, horen-snuit”

  • Proceedings Royal Society B

wo 17/07/2013 – 14:17 Luc De RoyAmerikaanse geleerden hebben een nieuwe soort dinosaurus beschreven, een erg ongewoon lid van de familie waartoe ook de triceratops behoort. Het gaat om een exemplaar met een erg grote neus en met de grootste horens die ooit bij een lid van de familie gevonden zijn.

De fossiele resten van de dinosaurus werden in 2006 gevonden in het Grand Staircase-Escalante Monument in de woestijn van Utah. Het heeft verschillende jaren geduurd om het fossiel te prepareren en te bestuderen en de resultaten zijn nu gepubliceerd in The Proceedings of the Royal Society B.

De nieuwe dinosaurus werd Nasutoceratops titusi genoemd, grote neus, hoorn-snuit; titusi verwijst naar de paleontoloog Alan Titus.

Proceedings Royal Society B

De rotsen waarin de fossielen gevonden werden dateren van zo’n 75 miljoen jaar geleden, de late Krijtperiode en de laatste bloeitijd van de dinosaurussen. Er werd een bijna volledige schedel gevonden, fragmenten van twee andere schedels en delen van de voorpoten.

Doctor Mark Loewen van de University of Utah en het Natural History Museum of Utah zei aan BBC News: “Deze dinosaurus blies ons compleet van onze sokken. We hadden nooit kunnen voorspellen dat hij er zo zou uitzien, – hij staat zo ver van de norm voor deze groep van dinosaurussen.”  Veel leden van de ceratops-famlilie hebben twee hoorns en een korte, neushoornachtige stomp op hun neus. In de plaats daarvan heeft Nasutoceratops een korte maar hoge neus met een boog op en erg lange hoorns.

Proceedings Royal Society B

“De hoorns zijn veruit de grootste van alle leden van deze groep van dinosaurussen”, zei Loewen, “ze buigen naar voren en naar opzij. Bovendien heeft hij ook de grootste neus van zijn groep.” Achteraan zijn kop had hij ook een gekartelde kraag.

Nasutoceratops was ook geen kleintje, hij zou zowat 2,5 ton gewogen hebben, en met zijn vreemde uiterlijk zou hij er angstaanjagend uitgezien hebben. Zoals alle leden van de ceratops-familie was hij evenwel een planteneter, die in de tropische, moerassige omgeving waar hij leefde, grote hoeveelheden planten verorberd zal hebben.

“Schatkamer”

De woestijn waar Nasutoceratops gevonden is, maakte deel uit van wat toen Laramidia was, een langgerekt eiland dat later met Appalachia Noord-Amerika zal vormen. Laramidia blijkt een ware schatkamer voor paleontologen.

Belga

In de buurt van de fossiele resten van Nautoceratops werden ook andere plantenetende soorten gevonden, waaronder twee andere soorten van gehoornde dinosaurussen en hadrosauriërs met een eendensnavel, wat er op wijst dat ze in staat waren samen te leven.

“Al deze dieren wogen verschillende tonnen… Je hebt een omgeving waarin al deze grote herbivoren wedijveren voor voedsel. We zijn niet echt zeker hoe je al deze dieren kunt onderhouden, maar je vindt ze wel allemaal tegelijkertijd in de rotsen”, zei Loewen. Hij voegde er nog aan toe dat er nog andere merkwaardige nieuwe soorten ontdekt waren.

Proceedings Royal Society B

(Proceedings of the Royal Society B)

Fossiel van broedende dino ontdekt

 22 februari 2012 1

Een bijzondere vondst in Mongolië.  Een moederdino gestorven en gefossiliseerd    terwijl ze op haar eieren broedde .De onderzoekers beschrijven hun ontdekking in het blad PLoS ONE. De dinosaurus in kwestie is afkomstig uit het geslacht Nemegtomaia. Toepasselijk genoeg betekent Nemegtomaia zoiets als: goede moeder van Nemegt (dit laatste is een verwijzing naar een geologische formatie waarin de dinosaurus is teruggevonden). Het geslacht bestaat uit dinosaurussen die zich op twee poten voortbewogen en al wel iets weghadden van de moderne vogels. De dinosaurus behoort tot de soort Nemegtomaia barsboldi.

Niet compleet
De gevonden dinosaurus is niet helemaal compleet. Er missen enkele delen. Waarschijnlijk is de dino nadat deze stierf door diverse roofdieren aangevreten. Maar wat er nog van over is, is toch tamelijk indrukwekkend. Zo laat het fossiel nog goed zien hoe de dinosaurus op de eieren zat. De poten bevonden zich hoogstwaarschijnlijk in het midden van een ring eieren en de armen had de dinosaurus om de bovenste delen van de eieren geslagen.

Een reconstructie van de dinosaurus op zijn nest. Afbeelding: via PLoS ONE – doi/10.1371/journal.pone.0031330.g003.
Ouder
De schedel van de dinosaurus is ongeveer 172 millimeter lang. Het dier moet ongeveer twee meter lang zijn geweest en ongeveer veertig kilo hebben gewogen. Het is zeer aannemelijk dat de dinosaurus één van de ouders van de jonge dino’s in de eieren was.

De gevonden eieren. Foto: via PLoS ONE – doi/10.1371/journal.pone.0031330.g010.
Het is niet helemaal duidelijk hoe de dinosaurus is gestorven. Het feit dat de dino al broedend stierf, wijst er echter op dat het dier een plotseling dood is gestorven. Mogelijk werd de dino overvallen door een zandstorm. Van de eieren is weinig overgebleven. Zo zijn er geen embryo’s of hele eieren meer aangetroffen. Wel kunnen de onderzoekers met zekerheid vaststellen dat de dino op zeker zeven eieren broedde. Deze eieren lagen wat dieper. Het is goed mogelijk dat zich op deze eieren nog een laag eieren bevond. De eieren moeten ongeveer vijf tot zes centimeter breed en veertien tot zestien centimeter lang zijn geweest.

A model of a Nodosaur dinosaur sits inside what is believed to be the fossil of a Nodosaur footprint. The footprint was found by Ray Stanford a local dinosaur hunter. (Credit: NASA/Goddard/Rebecca Roth)

http://www.sciencedaily.com/releases/2012/08/120821120324.htm

http://www.washingtonpost.com/national/health-science/dinosaur-age-meet-the-space-age/2012/08/17/76c176f4-e89a-11e1-8487-64e4b2a79ba8_story.html

Artist’s restoration of a pair of Nanshiungosaurus.

Choiniere, J., Forster, C., de Klerk, W. 2012. New information on Nqwebasaurus thwazi, a coelurosaurian theropod from the Earl Cretaceous Kirkwood Formation in South Africa. Journal of African Earth Sciences. 71-72: 1-17

click to enquire about this image

Nqwebasaurus thwazi            http://www.wildlifeartist.com.au/showimage.asp?code=F051

________________________________________________________________

  • Nyasasaurus” – nomen nudum; possibly non-dinosaurian (–> Archosauria)

-Dinosauria first appeared during the first period of the Mesozoic Era, the Triassic, and they prevailed throughout the rest of this Era. Currently, the birds are the only survivors of their lineage. Eoraptor, the proposed common ancester of dinosaurs is about 230 million years old, but recent (2012) studies of fossils like Nyasasaurus suggest the dino ancestors  may be older (250 million years old).

Meer afbeeldingen

Nyasasaurus parringtoni of Nyasasaurus alophos is een archosauriër, behorend tot de groep van de Dinosauriformes, die tijdens het Trias leefde in het gebied van het huidige Tanzania. Wellicht is het de oudste bekende dinosauriër.

Update –> Wikipedia

Nyasasaurus

Nyasasaurus parringtoni  Biol. Lett.-2013-Nesbitt- <– pdf  2012

nyasasaurus parringtoni

 

 

nyasasaurus in megamatrice

Nyasasaurus vs T alophos update

http://en.wikipedia.org/wiki/List_of_dinosaurs

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